Back in 1969, John Deere introduced a tractor designated the model "4000" It was essentially a stripped down 4020. It employed some components from previous models. (i.e., tranny/rear end parts from the 4010) No battery covers, no sheet metal covering seat suspention, less available options, and less over-all weight than the 4020. BUT. . . It had the SAME 96 HP 404 cu. in. diesel engine.
The purpose was twofold. One (probably being the MAIN reason) was to compete with the IH model 706. Bigger than Deere's 3020, and smaller than the 4020) It was quite a few bucks cheaper than a 4020. It filled a void in Deere's arsonal. #2 reason, and what Deere's advertising centered around was this. They claimed that a buyer who owned a 3020, and was ready to step up to more power, could buy a 4000, use the same implements as they were using with the 3020, and run FASTER in the field, at a lower throttle setting. That would get more work done, save a lot of cash on implements, and would be as productive as pulling bigger imlements at a slower speed, therefore, saving some money there. They also claimed field tests proved this concept (Gear up/throttle down) would save fuel.
The entire Deere line at that time was the "New Generation" tractors. They replaced the old 2 cylinder line. Their claim to fame was it's "HIGH HORSEPOWER TO WEIGHT RATIO". A typical 4010 was just a few pounds heavier than the 730 it replaced, but nearly double the available HP. They were both rated to pull a 5-bottom plow, only the 4010 would pull it almost twice as fast. Big fuel savings as well as higher productivity.
Also to consider, some engine manufacturers are better than others at fuel economy. And within those brands, some particular engine sizes are more economic.
The "standard" for the AG tractor industry has always been "The Nebraska Test". They rate HP hour per gallon for fuel economy. The big winners and big loosers in that area come from all sizes, big or small.