It's a vented system, and the fill plug doubles as the sole vent. It's this feature that makes the system self-priming. A clean fill/vent plug will intended to "inhale" as fluid is drawn from the sump (raise bucket/rear lift), and "exhale" as fluid flows back into the sump (lower bucket/rear lift). Yours sounds as if it's functioning properly, so air is apparently being drawn in elsewhere. Don't waste time starting on the pressure side of the pump either. As Ron points out, this type issue is typically found on the suction/return side.
//greg//