There are 2 numbers on circumference and have to be considered when running 4wd machines especially on hard surfaces like industrial applications where "Industrial R4s" are normally used...or cleaning poop out of your concrete floored barn/milking parlor. In tire discussions others have mentioned that they run their tire pressure on their equipment where the edge of the tread touches the pavement for a softer ride. It hasn't been mentioned, nor have I found where the load/tire pressure numbers for a given tire support the posted circumference, vs the "rolling circumference".
The point here is that once you deviate from the tractor designer's tire matching circumference, you mess with the tractors established gear ratio and as a result you put pressure on one of the drives and the weakest (obviously) is the front axle. In working my 2400 for a soft ride, I experimented with which axle was pushing and which was pulling with different types/sizes of rear tires run at varying pressures. As to which is best, I feel......gut feel, not verified by any broken parts, that you probably never will have the two perfectly matched and the "pusher" in the direction you normally run needs to be the rears with the heavier gear train. Had a friend put Monster Mudders on his Ford 4wd P/U and in a hard 4wd engaged turn, on concrete, the front would actually hop....how long that lasted I don't know but going to lunch one day he showed it to us.
Different numbers out there on recommended fills because you need some room for air. Max I've seen was 75% and I don't know if that's by volume, or height....didn't specify. HTH.