I had a forestry guy give me a lecture on the LONG life of a forest. It was quite interesting and I am sure what I am about to write has many missing points so take it as a starting point.
In a clearcut, weeds tend to come back first. Partly due to the Voracity, but in a bigger picture, the belief is that it attracts animals as a food source, which means fertilizer and new types of seeds.
In the PNW, the first trees up are generally Alders. They are finding a symbiotic relationship between Alders and Firs. Alders have a short life, so they will grow up, and provide protection from grazing animals for the Firs. First will eventually grow beyond the alders, choking them out.
In the fir lifecycle, I guess it is Firs First, then Red Cedar, then Hemlock. Of course if you are lucky you then get Sequoia / Redwood. This cycle is being listed over a 1000 years (meaning 100 years of Firs, followed by 150 years of cedar, then 150 years of hemlock, then.........
As for the thistle, outside of our berm, we just spray and mow. I have been told that for blackberry crossbow should be used spring and fall only, you are wasting it in a summer application. Not sure how it applies to thistle.
October 15 is my cutoff for winter spraying as I feel the plants are pretty much dormant by November 1.